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Handling Editor:
James McClintock, Birmingham, Alabama, USA

Reviewers:
C. Zhao and 2 anonymous referees

Acknowledgements:

We thank the Kuji City Fisheries Cooperative Association, the Oya-Motoyoshi, Shizugawa, and Onagawa-cho branches of the Miyagi Fisheries Cooperative Association, Kuji City, and Higashi-Matsushima City for granting permission to collect sea urchins during this field study. In addition, we are grateful to H. Kawajiri of the Kuji City Extension Center of Iwate University, Japan, for his support during the survey at Kuji; to M. Oshima and S. Kodama of Diving Stage Ariel for their support during the SCUBA diving survey at Kuji, Shizugawa Bay, and Shinchi; and to Tohoku Ryokka Kankyohozen and IDEA Consultants for their diving survey in Onagawa Bay. This study was financially supported by the TEAMS Fund of the Ministry of Education, Culture, Sports, Science, and Technology of Japan.